Flapping involves two stages: the down stroke, which provides the majority of the thrust, and the up stroke, which can also (depending on the bird’s wings) provide some thrust. At each up stroke the wing is slightly folded inwards to reduce the energetic cost of flapping wing flight.[8] Birds change the angle of attack continuously within a flap, as well as with speed.[9]Small birds often fly long distances using a technique in which short bursts of flapping are alternated with intervals in which the wings are folded against the body. This is a flight pattern known as “bounding” or “flap bounding” flight.[10] When the bird’s wings are folded, its trajectory is primarily ballistic, with a small amount of body lift.[2] The flight pattern is believed to decrease the energy required by reducing the aerodynamic drag during the ballistic part of the trajectory,[11] and to increase the efficiency of muscle use.[12][13].

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